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Because the disulfide bridge (a cyclic disulfide) can undergo redox reactions, during the reactions catalyzed by the pyruvate dehydrogenase complex it is first reduced to dihydrolipoamide, a dithiol (the reduced form of the prosthetic group), and then reoxidized to the cyclic form

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(PubMed) Sawicka AK, Hartmane D, Lipinska P, Wojtowicz E, Lysiak-Szydlowska W, Olek RA