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ATF-4 siRNA transfection The siRNA used to downregulate human ATF-4 and the non-targeting siRNA control were purchased from Dharmacon (siGENOME SMART pool siRNA)

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Heights of the different zones of chondrocytes were measured parallel to the chondrocyte columns using the MacBAS version 2.5 program (Koshin Graphic Systems, Sapporo, Japan)

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