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Ser is a substrate for mitochondrial onecarbon metabolism, which generates NADPH while converting Ser + tetrahydrofolate (THF) into glycine (Gly) + THF + formate through the activity of serine hydroxymethyl transferase 2 (SHMT2), methyleneTHF dehydrogenase (MTHFD) 2, and MTHFD1L

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Additionally, the IMM is equipped with numerous transport systems that regulate the movement of metabolites across the membrane, ensuring the efficient production and distribution of energy within the cell (Picard and Shirihai, 2022)